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Stylops melittae





Subspecies
The genus Stylops previously contained a large number of species which were believed to be highly specialized for a specific host bee species. However, Kinzelbach (1978) argued that the criteria for species status in Stylops are unclear and require much more work. In particular, the "species" are much less host bee species specific than previously thought, thus calling into question many of the
described Stylops "species". Kinzelbach (1978) therefore united all Stylops "species" under Stylops melittae, some as valid subspecies, some as synonyms. On the other hand, this renders the currently recognized species Stylops melittae a very variable species that clearly consists of several natural subunits that may be subspecies or even separate species. Unless more work is done on the taxonomy of this species, it is impossible to give a confident number of subspecies or separate species. Stylops melittae is therefore currently best regarded as "Stylops melittae-species complex" that requires substantial additional work to recognize the biological species and subspecies contained in it. The subspecies recognized by Kinzelbach (1978) are given below with their synonyms.

Stylops melittae melittae Kirby, 1802
     Stylops melittae Kirby, 1802
     Stylops melittae melittae Kirby, 1802
     Stylops melittae kirbii Leach, 1817
     Stylops kirbii Leach, 1817
     Stylops melittae kirbyi Leach, 1817 (justified emendation)
     Stylops kirbyi Leach, 1817 (justified emendation)
     Stylops melittae haworthi Stephen, 1929
     Stylops haworthi Stephen, 1929
     Stylops mellitae auct. (a misspelling)

Stylops melittae alfkeni Hofeneder, 1939
     Stylops alfkeni Hofeneder, 1939
     Stylops melittae alfkeni Hofeneder, 1939

Stylops melittae analis Perkins, 1918
     Stylops analis Perkins, 1918
     Stylops melittae analis Perkins, 1918

Stylops melittae andrenaphilus Luna de Carvalho, 1974
    Stylops andrenaphilus Luna de Carvalho, 1974
    Stylops melittae andrenaphilus Luna de Carvalho, 1974

Stylops melittae bimaculatae Perkins, 1918
     Stylops bimaculatae Perkins, 1918
     Stylops melittae bimaculatae Perkins, 1918

Stylops melittae borcherti Luna de Carvalho, 1974
     Stylops borchertai
Luna de Carvalho, 1974
     Stylops borcherti Luna de Carvalho, 1974 (justified emendation)
     Stylops melittae borcherti Luna de Carvalho, 1974 (justified emendation)

Stylops melittae championi Pierce, 1918
     Stylops championi Pierce, 1918
     Stylops melittae championi Pierce, 1918

Stylops melittae dalei Curtis, 1828
     Stylops melittae dalii Curtis, 1828
     Stylops dalii Curtis, 1828
     Stylops dalei Curtis, 1828 (justified emendation)
     Stylops melittae dalei Curtis, 1828 (justified emendation)

Stylops melittae deserticola Medvedev, 1970
     Stylops deserticola Medvedev, 1970
     Stylops melittae deserticola Medvedev, 1970
     Stylops desertorum auct. (a misspelling or unjustified emendation)

Stylops melittae dinizi Luna de Carvalho, 1974
    
Stylops dinizai Luna de Carvalho, 1974
     Stylops dinizi Luna de Carvalho, 1974 (justified emendation)
     Stylops melittae dinizi Luna de Carvalho, 1974 (justified emendation)

Stylops melittae dominiquei Pierce, 1909
     Stylops dominiquei Pierce, 1909
     Stylops melittae dominiquei Pierce, 1909

Stylops melittae duriensis Luna de Carvalho, 1974
     Stylops duriensis Luna de Carvalho, 1974
     Stylops melittae duriensis Luna de Carvalho, 1974

Stylops melittae esteponensis Luna de Carvalho, 1974
     Stylops esteponensis Luna de Carvalho, 1974
     Stylops melittae esteponensis Luna de Carvalho, 1974

Stylops melittae giganteus Luna de Carvalho, 1974
     Stylops giganteus Luna de Carvalho, 1974
     Stylops melittae giganteus Luna de Carvalho, 1974

Stylops melittae gwynanae Noskiewicz et Poluszynski, 1927
     Stylops gwynanae Noskiewicz et Poluszynski, 1927
     Stylops melittae gwynanae Noskiewicz et Poluszynski, 1927

Stylops melittae hammellae Perkins, 1918
     Stylops hammellae Perkins, 1918
     Stylops melittae hammellae Perkins, 1918

Stylops melittae ibericus Luna de Carvalho, 1969
     Stylops ibericus Luna de Carvalho, 1969
     Stylops melittae ibericus Luna de Carvalho, 1969

Stylops melittae kinzelbachi Luna de Carvalho, 1974
    
Stylops kinzelbachai Luna de Carvalho, 1974
     Stylops kinzelbachi Luna de Carvalho, 1974 (justified emendation)
     Stylops melittae kinzelbachi Luna de Carvalho, 1974 (justified emendation)

Stylops melittae liliputanus Luna de Carvalho, 1974
     Stylops liliputanus Luna de Carvalho, 1974
     Stylops melittae liliputanus Luna de Carvalho, 1974
     Stylops melittae lusohispanicus Luna de Carvalho, 1974 (unclear synonymy)
     Stylops lusohispanicus Luna de Carvalho, 1974 (unclear synonymy)

Stylops melittae madrilensis Luna de Carvalho, 1974
     Stylops madrilensis Luna de Carvalho, 1974
     Stylops melittae madrilensis Luna de Carvalho, 1974

Stylops melittae maxillaris Pasteels, 1949
     Stylops maxillaris Pasteels, 1949
     Stylops melittae maxillaris Pasteels, 1949

Stylops melittae moniliaphagus Luna de Carvalho, 1974
     Stylops moniliaphagus Luna de Carvalho, 1974
     Stylops melittae moniliaphagus Luna de Carvalho, 1974

Stylops melittae nassonowi Pierce, 1909
     Stylops melittae Nassonov, 1893 (nec Kirby, 1802)
     Stylops nassonowi Pierce, 1909
     Stylops melittae nassonowi Pierce, 1909
     Stylops pilipedis Pierce, 1911
     Stylops melittae pilipedis Pierce, 1911
     Stylops flavipedis Hofeneder, 1923 (unclear synonymy)
     Stylops melittae flavipedis Hofeneder, 1923 (unclear synonymy)
     Stylops obenbergeri Ogloblin, 1923
     Stylops melittae obenbergeri Ogloblin, 1923
     Stylops ruthenicus Schkaff, 1925 (synonym of flavipedis?)
     Stylops melittae ruthenicus Schkaff, 1925 (synonym of flavipedis?)
     Stylops nassonovi auct. (a misspelling)

Stylops melittae nevinsoni Perkins, 1918
     Stylops nevinsoni Perkins, 1918
     Stylops melittae nevinsoni Perkins, 1918
     Stylops praecocis Noskiewicz et Poluszynski, 1927
     Stylops melittae praecocis Noskiewicz et Poluszynski, 1927

Stylops melittae nitidae Pasteels, 1954
     Stylops nitidae Pasteels, 1954
     Stylops melittae nitidae Pasteels, 1954

Stylops melittae nitidiusculae Poluszynski, 1927
     Stylops nitidiusculae Poluszynski, 1927
     Stylops melittae nitidiusculae Poluszynski, 1927

Stylops melittae nycthemerae Noskiewicz et Poluszynski, 1927
     Stylops nycthemerae Noskiewicz et Poluszynski, 1927
     Stylops melittae nycthemerae Noskiewicz et Poluszynski, 1927

Stylops melittae obsoletus Luna de Carvalho, 1974
     Stylops obsoletus Luna de Carvalho, 1974
     Stylops melittae obsoletus Luna de Carvalho, 1974

Stylops melittae ovinae Noskiewicz et Poluszynski, 1927
     Stylops melittae ovinae Noskiewicz et Poluszynski, 1927
     Stylops melittae ovinae Noskiewicz et Poluszynski, 1927
     Stylops melittae krygeri Pierce, 1918 (unclear synonymy)
     Stylops krygeri Pierce, 1918 (unclear synonymy)
     Stylops melittae muelleri Borchert, 1971
     Stylops muelleri Borchert, 1971

Stylops melittae paracuellus Luna de Carvalho, 1974
     Stylops paracuellus Luna de Carvalho, 1974
     Stylops melittae paracuellus Luna de Carvalho, 1974

Stylops melittae parvulae Noskiewicz et Poluszynski, 1927
     Stylops parvulae Noskiewicz et Poluszynski, 1927
     Stylops melittae parvulae Noskiewicz et Poluszynski, 1927

Stylops melittae pasteelsi Luna de Carvalho, 1974
    
Stylops pasteelsai Luna de Carvalho, 1974
     Stylops pasteelsi Luna de Carvalho, 1974 (justified emendation)
     Stylops melittae pasteelsi Luna de Carvalho, 1974 (justified emendation)

Stylops melittae perkinsi Pasteels, 1949
     Stylops perkinsi Pasteels, 1949
     Stylops melittae perkinsi Pasteels, 1949
     Stylops spencii Perkins, 1918 (nec Pickering, 1836)
     Stylops melittae spencii Perkins, 1918 (nec Pickering, 1836)

Stylops melittae salamancanus Luna de Carvalho, 1974
     Stylops salamancanus Luna de Carvalho, 1974
     Stylops melittae salamancanus Luna de Carvalho, 1974

Stylops melittae savignyi Hofeneder, 1924
     Stylops savignyi Hofeneder, 1924
     Stylops melittae savignyi Hofeneder, 1924

Stylops melittae spencei Pickering, 1836
     Stylops spencii Pickering, 1836
     Stylops melittae spencii Pickering, 1836
     Stylops melittae spencei Pickering, 1836 (justified emendation)
     Stylops spencei Pickering, 1836 (justified emendation)
     Stylops melittae aterrimus Newport, 1851
     Stylops aterrimus Newport, 1851
     Stylops melittae niger von Beneden, 1875
     Stylops niger von Beneden, 1875
     Stylops melittae trimmeranae Smith, 1957
     Stylops trimmeranae Smith, 1957

Stylops melittae spretae Perkins, 1918
     Stylops spreta Perkins, 1918
     Stylops spretae Perkins, 1918 (justified emendation)
     Stylops melittae spretae Perkins, 1918 (justified emendation)
     Stylops risleri Kinzelbach, 1967
     Stylops melittae risleri Kinzelbach, 1967

Stylops melittae thwaitei Saunders, 1872
     Stylops thwaitei Saunders, 1872
     Stylops melittae thwaitei Saunders, 1872
     Stylops wilkellae Perkins, 1918
     Stylops melittae wilkellae Perkins, 1918
     Stylops melittae albofasciatae Günther, 1957
     Stylops albofasciatae Günther, 1957

Stylops melittae transversus Pasteels, 1949
     Stylops transversus Pasteels, 1949
     Stylops melittae transversus Pasteels, 1949

Stylops melittae ventricosae Pierce, 1909
     Stylops ventricosae Pierce, 1909
     Stylops melittae ventricosae Pierce, 1909

Stylops melittae warnckei Luna de Carvalho, 1974
     Stylops warnckeai Luna de Carvalho, 1974
     Stylops warnckei Luna de Carvalho, 1974 (justified emendation)
     Stylops melittae warnckei Luna de Carvalho, 1974 (justified emendation)



Synonyms
See above in the chapter Subspecies.



Distribution
Many of the subspecies are known from a limited set of localities only. This does not necessarily indicate a restricted geographical distribution of these subspecies, rather it reflects our current ignorance of the biology and biogeography of Stylops melittae. The following subspecies have been recorded in Germany:
Stylops melittae melittae, Stylops melittae alfkeni, Stylops melittae andrenaphilus, Stylops melittae bimaculatae, Stylops melittae dalei, Stylops melittae nassonowi (also under its synonym Stylops melittae flavipedis), Stylops melittae gwynanae, Stylops melittae hammellae, Stylops melittae maxillaris, Stylops melittae ovinae (under its synonym Stylops melittae muelleri), Stylops melittae nitidae, Stylops melittae nitidiusculae, Stylops melittae nycthemerae, Stylops melittae parvulae, Stylops melittae perkinsi, Stylops melittae nevinsoni (under its synonym Stylops melittae praecocis), Stylops melittae thwaitei, Stylops melittae transversus (also under its synonym Stylops melittae wilkellae). Taken together Stylops melittae is recorded in Germany from all Bundesländer, except for Bayern. The lack of bavarian records is unusual and urgently needs re-investigation.











This page has been updated on March 29, 2009
This site is online since May 31, 2005
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