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Stylops melittae
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Subspecies
The genus Stylops previously contained a large
number of species which were believed to be highly specialized for a
specific
host bee species. However, Kinzelbach (1978) argued that the criteria
for species status in Stylops are unclear and require much more work.
In particular, the "species" are much less host bee species specific
than
previously thought, thus calling into question many of the described Stylops "species". Kinzelbach (1978) therefore
united all Stylops "species" under Stylops melittae, some as valid
subspecies, some as synonyms. On the other hand,
this renders the currently recognized species Stylops melittae a very
variable
species that clearly consists of several natural subunits that may be
subspecies or even separate species. Unless more work is done on the
taxonomy of this species, it is impossible to give a confident number
of
subspecies or separate species. Stylops melittae is therefore currently
best regarded as "Stylops melittae-species complex" that requires
substantial additional work to recognize the biological species and
subspecies contained in it. The subspecies recognized by Kinzelbach
(1978) are given below with their synonyms.
Stylops melittae melittae Kirby, 1802
Stylops melittae Kirby, 1802
Stylops melittae melittae Kirby, 1802
Stylops melittae kirbii Leach, 1817
Stylops kirbii Leach, 1817
Stylops melittae kirbyi Leach, 1817 (justified emendation)
Stylops kirbyi Leach, 1817 (justified emendation)
Stylops melittae haworthi Stephen, 1929
Stylops haworthi Stephen, 1929
Stylops mellitae auct. (a misspelling)
Stylops
melittae alfkeni Hofeneder, 1939
Stylops alfkeni Hofeneder, 1939
Stylops melittae alfkeni Hofeneder, 1939
Stylops
melittae analis Perkins, 1918
Stylops analis Perkins, 1918
Stylops melittae analis Perkins, 1918
Stylops melittae andrenaphilus Luna de
Carvalho, 1974
Stylops andrenaphilus Luna de Carvalho, 1974
Stylops melittae andrenaphilus Luna de Carvalho, 1974
Stylops
melittae bimaculatae Perkins, 1918
Stylops bimaculatae Perkins, 1918
Stylops melittae bimaculatae Perkins, 1918
Stylops
melittae borcherti Luna de Carvalho, 1974
Stylops borchertai Luna de Carvalho, 1974
Stylops borcherti Luna de Carvalho, 1974 (justified emendation)
Stylops melittae borcherti Luna de Carvalho, 1974 (justified emendation)
Stylops
melittae championi Pierce, 1918
Stylops championi Pierce, 1918
Stylops melittae championi Pierce, 1918
Stylops
melittae dalei Curtis, 1828
Stylops melittae dalii Curtis, 1828
Stylops dalii Curtis, 1828
Stylops dalei Curtis, 1828 (justified emendation)
Stylops melittae dalei Curtis, 1828 (justified emendation)
Stylops
melittae deserticola Medvedev, 1970
Stylops deserticola Medvedev, 1970
Stylops melittae deserticola Medvedev, 1970
Stylops desertorum auct. (a misspelling or
unjustified emendation)
Stylops
melittae dinizi Luna de Carvalho, 1974
Stylops dinizai Luna de Carvalho, 1974
Stylops dinizi Luna de Carvalho, 1974 (justified emendation)
Stylops melittae dinizi Luna de Carvalho, 1974 (justified emendation)
Stylops
melittae dominiquei Pierce, 1909
Stylops dominiquei Pierce, 1909
Stylops melittae dominiquei Pierce, 1909
Stylops
melittae duriensis Luna de Carvalho, 1974
Stylops duriensis Luna de Carvalho, 1974
Stylops melittae duriensis Luna de Carvalho, 1974
Stylops
melittae esteponensis Luna de Carvalho, 1974
Stylops esteponensis Luna de Carvalho, 1974
Stylops melittae esteponensis Luna de Carvalho, 1974
Stylops melittae giganteus Luna de
Carvalho, 1974
Stylops giganteus Luna de Carvalho, 1974
Stylops melittae giganteus Luna de Carvalho, 1974
Stylops
melittae gwynanae Noskiewicz et Poluszynski, 1927
Stylops gwynanae Noskiewicz et Poluszynski, 1927
Stylops melittae gwynanae Noskiewicz et Poluszynski, 1927
Stylops
melittae hammellae Perkins, 1918
Stylops hammellae Perkins, 1918
Stylops melittae hammellae Perkins, 1918
Stylops
melittae ibericus Luna de Carvalho, 1969
Stylops ibericus Luna de Carvalho, 1969
Stylops melittae ibericus Luna de Carvalho, 1969
Stylops
melittae kinzelbachi Luna de Carvalho, 1974
Stylops kinzelbachai Luna de Carvalho, 1974
Stylops kinzelbachi Luna de Carvalho, 1974 (justified emendation)
Stylops melittae kinzelbachi Luna de Carvalho, 1974 (justified
emendation)
Stylops
melittae liliputanus Luna de Carvalho, 1974
Stylops liliputanus Luna de Carvalho, 1974
Stylops melittae liliputanus Luna de Carvalho, 1974
Stylops melittae lusohispanicus Luna de Carvalho, 1974 (unclear synonymy)
Stylops lusohispanicus Luna de Carvalho, 1974 (unclear synonymy)
Stylops
melittae madrilensis Luna de Carvalho, 1974
Stylops madrilensis Luna de Carvalho, 1974
Stylops melittae madrilensis Luna de Carvalho, 1974
Stylops
melittae maxillaris Pasteels, 1949
Stylops maxillaris Pasteels, 1949
Stylops melittae maxillaris Pasteels, 1949
Stylops
melittae moniliaphagus Luna de Carvalho, 1974
Stylops moniliaphagus Luna de Carvalho, 1974
Stylops melittae moniliaphagus Luna de Carvalho, 1974
Stylops
melittae nassonowi Pierce, 1909
Stylops melittae Nassonov, 1893 (nec Kirby, 1802)
Stylops nassonowi Pierce, 1909
Stylops melittae nassonowi Pierce, 1909
Stylops pilipedis Pierce, 1911
Stylops melittae pilipedis Pierce, 1911
Stylops flavipedis Hofeneder, 1923 (unclear synonymy)
Stylops melittae flavipedis Hofeneder, 1923 (unclear synonymy)
Stylops obenbergeri Ogloblin, 1923
Stylops melittae obenbergeri Ogloblin, 1923
Stylops ruthenicus Schkaff, 1925 (synonym of flavipedis?)
Stylops melittae ruthenicus Schkaff, 1925 (synonym of flavipedis?)
Stylops nassonovi auct. (a misspelling)
Stylops
melittae nevinsoni Perkins, 1918
Stylops nevinsoni Perkins, 1918
Stylops melittae nevinsoni Perkins, 1918
Stylops praecocis Noskiewicz et Poluszynski, 1927
Stylops melittae praecocis Noskiewicz et Poluszynski, 1927
Stylops
melittae nitidae Pasteels, 1954
Stylops nitidae Pasteels, 1954
Stylops melittae nitidae Pasteels, 1954
Stylops
melittae nitidiusculae Poluszynski, 1927
Stylops nitidiusculae Poluszynski, 1927
Stylops melittae nitidiusculae Poluszynski, 1927
Stylops
melittae nycthemerae Noskiewicz et Poluszynski, 1927
Stylops nycthemerae Noskiewicz et Poluszynski, 1927
Stylops melittae nycthemerae Noskiewicz et Poluszynski, 1927
Stylops
melittae obsoletus Luna de Carvalho, 1974
Stylops obsoletus Luna de Carvalho, 1974
Stylops melittae obsoletus Luna de Carvalho, 1974
Stylops melittae ovinae Noskiewicz et Poluszynski, 1927
Stylops melittae ovinae Noskiewicz et Poluszynski, 1927
Stylops melittae ovinae Noskiewicz et Poluszynski, 1927
Stylops melittae krygeri Pierce, 1918 (unclear synonymy)
Stylops krygeri Pierce, 1918 (unclear synonymy)
Stylops melittae muelleri Borchert, 1971
Stylops muelleri Borchert, 1971
Stylops
melittae paracuellus Luna de Carvalho, 1974
Stylops paracuellus Luna de Carvalho, 1974
Stylops melittae paracuellus Luna de Carvalho, 1974
Stylops
melittae parvulae Noskiewicz et Poluszynski, 1927
Stylops parvulae Noskiewicz et Poluszynski, 1927
Stylops melittae parvulae Noskiewicz et Poluszynski, 1927
Stylops
melittae pasteelsi Luna de Carvalho, 1974
Stylops pasteelsai Luna de Carvalho, 1974
Stylops pasteelsi Luna de Carvalho, 1974 (justified emendation)
Stylops melittae pasteelsi Luna de Carvalho, 1974 (justified emendation)
Stylops
melittae perkinsi Pasteels, 1949
Stylops perkinsi Pasteels, 1949
Stylops melittae perkinsi Pasteels, 1949
Stylops spencii Perkins, 1918 (nec Pickering, 1836)
Stylops melittae spencii Perkins, 1918 (nec Pickering, 1836)
Stylops
melittae salamancanus Luna de Carvalho, 1974
Stylops salamancanus Luna de Carvalho, 1974
Stylops melittae salamancanus Luna de Carvalho, 1974
Stylops
melittae savignyi Hofeneder, 1924
Stylops savignyi Hofeneder, 1924
Stylops melittae savignyi Hofeneder, 1924
Stylops melittae spencei Pickering, 1836
Stylops spencii Pickering, 1836
Stylops melittae spencii Pickering, 1836
Stylops melittae spencei Pickering, 1836 (justified emendation)
Stylops spencei Pickering, 1836 (justified emendation)
Stylops melittae aterrimus Newport, 1851
Stylops aterrimus Newport, 1851
Stylops melittae niger von Beneden, 1875
Stylops niger von Beneden, 1875
Stylops melittae trimmeranae Smith, 1957
Stylops trimmeranae Smith, 1957
Stylops melittae spretae Perkins, 1918
Stylops spreta Perkins, 1918
Stylops spretae Perkins, 1918 (justified emendation)
Stylops melittae spretae Perkins, 1918 (justified emendation)
Stylops risleri Kinzelbach, 1967
Stylops melittae risleri Kinzelbach, 1967
Stylops
melittae thwaitei Saunders, 1872
Stylops thwaitei Saunders, 1872
Stylops melittae thwaitei Saunders, 1872
Stylops wilkellae Perkins, 1918
Stylops melittae wilkellae Perkins, 1918
Stylops melittae albofasciatae Günther, 1957
Stylops albofasciatae Günther, 1957
Stylops
melittae transversus Pasteels, 1949
Stylops transversus Pasteels, 1949
Stylops melittae transversus Pasteels, 1949
Stylops
melittae ventricosae Pierce, 1909
Stylops ventricosae Pierce, 1909
Stylops melittae ventricosae Pierce, 1909
Stylops
melittae warnckei Luna de Carvalho, 1974
Stylops warnckeai Luna de Carvalho, 1974
Stylops warnckei Luna de Carvalho, 1974 (justified emendation)
Stylops melittae warnckei Luna de Carvalho, 1974 (justified emendation)
Synonyms
See above in the chapter Subspecies.
Distribution
Many of the subspecies are known from a limited set
of localities only. This does not necessarily indicate a restricted
geographical distribution of these subspecies, rather it reflects our
current ignorance of the biology and biogeography of Stylops melittae.
The following subspecies have been recorded in Germany: Stylops melittae melittae, Stylops melittae alfkeni, Stylops melittae andrenaphilus, Stylops melittae bimaculatae, Stylops melittae dalei, Stylops melittae nassonowi (also under its
synonym Stylops
melittae flavipedis), Stylops melittae gwynanae, Stylops melittae hammellae, Stylops melittae maxillaris, Stylops melittae ovinae (under its synonym
Stylops
melittae muelleri), Stylops melittae nitidae, Stylops melittae nitidiusculae, Stylops melittae nycthemerae, Stylops melittae parvulae, Stylops melittae perkinsi, Stylops melittae nevinsoni (under its
synonym Stylops
melittae praecocis), Stylops melittae thwaitei, Stylops melittae transversus (also under
its synonym Stylops
melittae wilkellae). Taken together Stylops melittae is recorded in
Germany from all Bundesländer, except for Bayern. The lack of
bavarian records is unusual and urgently needs re-investigation.
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This page has been
updated on March 29, 2009
This site is online since May 31, 2005
Copyright © by Nikola-Michael Prpic. All rights reserved.
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