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Bufo calamita
Natterjack Toad
Kreuzkröte
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Subspecies
No subspecies are recognized. The status of Bufo calamita alpestris Koch, 1872 is debated; it is currently placed in synonymy (see below).


Synonyms
Bufo calamita Laurenti, 1768
Epidalea calamita (Laurenti, 1768)
Bufo viridis calamita (Laurenti, 1768)
Rana foetidissima Hermann, 1783
Rana portentosa Blumenbach, 1788
Bufo portentosa (Blumenbach, 1788)
Rana ecaudata Razoumowsky, 1789
Bufo cruciatus Schneider, 1799
Rana mephitica Shaw, 1802
Bufo cursor Daudin, 1803
Bufo calamita alpestris Koch, 1872
Rana fetidissima Boulenger, 1898 (unjustified emendation)

Bufo calamita has been placed in its own genus Epidalea by Frost et al. 2006. These authors argue that the genus Bufo in its traditional form is paraphyletic, because, in their analysis, a small number of "non-Bufo" species are grouped together with the Bufo species, and a couple of canonical Bufo species do not group with the large bulk of Bufo species.
This is not followed here for two reasons:
(1) The phylogeny of the Bufonidae is still unclear as recent studies do not agree on a common phylogenetic pattern (e.g. Stoeck et al. 2006; Pramuk et al. 2008). Most importantly, in the analysis by Pramuk et al. 2008 the Eurasian species appear to form a well-defined monophyletic group. This suggests that the Eurasian species (including the type species of Bufo) form a closely related assemblage sharing a common biogeographical history.
(2) The main duty of zoological nomenclature is to ensure stability as far as possible. Thus, even if one accepts the phylogeny in Frost et al. 2006, the easiest measure to avoid paraphyletic groups and to change as little as possible would be to retain Bufo sensu lato, include those few species that cluster within it also into Bufo and distribute the few Bufo species that do not cluster with the bulk of Bufo to other genera. The authors, however, erect or resurrect a large number of genera and thus split Bufo into numerous new or resurrected genera, although they state themselves that their results are just a first step towards a better understanding of Bufonidae phylogeny and they expect it to change as more work is done.
Therefore I do not adopt the splitting of the Eurasian Bufo into numerous genera until more data on the phylogenetic interrelationships of the Bufonidae is available.


Identification
No information has been entered yet.


Distribution
Germany, Switzerland, Belgium, Netherlands, Poland, Czech Republic, Slovakia, western part of Russia, Iberian peninsula, France, Great Britain, Denmark, Sweden (southern part), Latvia, Lithuania.


Biology
No information has been entered yet.


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This site is online since May 31, 2005
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