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Familia Atypidae
General information
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Purse-web spiders
are relatively large and very robust spiders.
Their general appearance is similar to tropical
bird spiders and indeed they belong to the same
group of basal spiders, the Mygalomorphae.
Characteristics of the Mygalomorphae are the
possession of two pairs of book lungs (higher
spiders have only one pair of book lungs, while
the second breathing organ is a tracheal system)
and the orthognathous chelicerae (i.e. the claws
project forward and not sideways as in
labidognathous spiders). The Atypidae have
extraordinarily strong and large chelicerae and
a heavily sclerotized prosomal shield that is
reminiscent of the carapace of crabs. The
species that occur in Germany are dark brown or
black and about 10-15 mm long (the males are a
bit smaller).
Purse-web spiders build tubes in loose soil that
are lined on the inside with silk. This
subterranean portion of the tube can lead up to
a meter deep in the ground. The silk lining
protudes from the tube and forms a silk sack on
the surface. This sack is camouflaged by the
spider with detritus, plant parts and earth
crumbs and thus resembles a plant root or
decaying plant parts. When prey crawls or lands
on this camouflaged external portion of the silk
tube, the spider in the subterranean portion
darts out and grabs the prey through the silk
sack. Then the spider rips a hole in the
external silk sack, wraps the prey in it and
carries the immobilized prey into the
subterranean part of the tube. Later the spider
returns to the surface and repairs the hole in
the external silk sack and also refits the
camouflage of the sack.
Dispersal only takes place during nymphal stages
in the springtime (see also below); soon after
this dispersal phase the spiderlings build their
own tubes in which they spend most of their
life. In Central Europe, the juveniles take 4
years to reach maturity in late July to
September (depending on the species) of the
fourth year. Females stay in their tubes after
reaching sexual maturity, but the mature males
leave their tubes and search for the tubes of
females. By drumming a specific rhythmic pattern
with their legs and pedipalps on the external
silk sack of the female the males are able to
enter the tube of the female without being
mistaken for prey. The pair then mates and lives
together in the same tube until the onset of
winter. Then the male dies or is eaten by the
female. The female then produces an egg sac and
the young spiders hatch in late summer. These
juveniles remain with the mother over winter
until they leave the tube and spread to new
areas in the springtime of the following year.
Thus, the time span between mating and spreading
of the juveniles is about 18 months. The
juveniles mainly spread by "balooning", i.e.
they produce long silk threads that catch wind
gusts and take the spiderlings to air. The
females are very long-lived and can mate several
times in their live. They can live up to 10
years, whereas the males do not survive their
maturity for very long and, thus, rarely get
much older than 4 years.
The German species prefer sandy or limy soil
often along forest edges with dense vegetation.
The animals need sunny, dry and warm slopes to
build their tubes, but the steepness of the
slopes differs from species to species. Some
species can use only slightly oblique surfaces,
while others require steep slopes of well over
45°.
If an Atypidae spider is threatened or teased
out of its tube, it shows a special defensive
behavior. The spider lifts its prosoma and first
leg pair and displays its huge chelicerae with
its claws. This view is impressive enough to
intimidate virtually every predator.
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This page has
been updated on December 17, 2012
This site is online since May 31, 2005
Copyright © by Nikola-Michael Prpic. All
rights reserved.
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