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Diplocentria rectangulata


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Subspecies
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Synonyms
Tmeticus rectangulatus Emerton, 1915 (p. 137)
Microcentria rectangulata (Emerton, 1915)
Diplocentria rectangulata (Emerton, 1915)
Parerigone rectangulatus (Emerton, 1915)
Type locality: "Mt. Mansfield, Vt., and Brunswick, Me.".
Microcentria pusilla Schenkel, 1925 (p. 297)
Type locality: not given. Switzerland (as inferred from title).


Note: the taxon Savignia foveata, though synonymized by Wiehle (1960), is in fact not a synonym of the present species (see Erigonoplus foveatus for more information).

Note: there is confusion concerning the identity of two taxa described by Emerton for both of which he used the species epithet "rectangulatus/-a". Emerton (1913) described a new species Microneta rectangulata, and Emerton (1915) described another new species Tmeticus rectangulatus. The World Spider Catalogue (Platnik, 2000ff., accessed on January 18, 2014) lists Microneta rectangulata (sub Oreonetides) and Tmeticus rectangulatus (sub Diplocentria) as separate species, but then also lists the younger name as a synonym of the older name. The latter is clearly a mistake; both taxa are separate species, which is already obvious from their original description:

Emerton 1913, p. 217
"Microneta rectangulata. new. 2 mm. long, light orange brown with dark gray abdomen. The mandibles are thickened at the base as in M. viaria, and narrow at the tip with a small tooth on the front just below the thickened part. The palpi have the tarsus comparatively narrow with a very small spur at the base. The tarsal hook is bent at a right angle, the outer portion thickened at the corner, but ending in a thin, wide tip. Figs. 5, 5a, 5b. Pl. II. Readville, Mass., on a fence in the autumn flight, Nov. 5, 1911".

Emerton 1915, p. 137
"Tmeticus rectangulatus. new sp. A translucent spider 1.5 mm. long, resembling T. entomologicus from Tyngsboro, Mass., 1911; T. digitatus, Em., from Ithaca, N. Y., Journal N. Y. Ent. Soc, 1914, and T. acummatus, Em., from New Jersey, Bulletin Am. Museum, N. Y., 1913. It differs from these species in the tibia of the male palpus, which is truncate and has three small teeth across the end. Fig. 4. Pl. I. One male each from Mt. Mansfield, Vt., and Brunswick, Me.".

The original illustrations (shown below) also show that Microneta rectangulata and Tmeticus rectangulatus are different species:


These images have been copied from the original species descriptions by Emerton (1913), Emerton (1915) and Schenkel (1925) and were rotated and scaled to achieve maximum comparability. They show the distal end of the male pedipalp, with the bulb apparatus and the cymbium. The shape of the cymbium clearly differs between Microneta rectangulata and Tmeticus rectangulatus (note the lateral outgrowth in Microneta rectangulata!). In addition, the shape of the paracymbium differs, it is a angular hook in Microneta rectangulata, whereas it is a rounded hook in Tmeticus rectangulatus. In conclusion, Tmeticus rectangulatus is not a synonym of Microneta rectangulata; both are differentent species. By contrast, the original figures of Tmeticus rectangulatus and Microcentria pusilla show that both are very similar and that the synonymization of both names is justified. The shape of the paracymbium is a rounded hook in both cases and the characteristic scoop-shape of the cymbium is identical in both taxa. In addition, the distal end of the tibia shows three protrusions (the third cannot be seen in Schenkel´s figure, because the pedipalp is viewed from a slighly different angle than the one in Emerton´s figure, but is described in the text of the original description).


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