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Archiearis parthenias
Orange Underwing 
Birken-Jungfernkind 
Subspecies
(1) Archiearis parthenias parthenias (Linnaeus, 1761)
(2) Archiearis parthenias bella (Inoue, 1955)
(3) Archiearis parthenias elegans (Inoue, 1955)
(4) Archiearis parthenias hilara (Sawamoto, 1937)
(5) Archiearis parthenias lapponica (Rangnow, 1935)
(6) Archiearis parthenias sajana (Prout (in Seitz), 1912)

The status of the taxon lapponica is unclear. Hausmann (2001) expresses doubts about the identity of the type specimen and lists the taxon as a doubtful synonym of the nominate subspecies. Only the nominate subspecies occurs in Germany.
Synonyms
Synonymy of Archiearis parthenias parthenias:

Phalaena parthenias Linnaeus, 1761
Archiearis parthenias (Linnaeus, 1761)
Archiearis parthenias parthenias (Linnaeus, 1761)
Brephos parthenias (Linnaeus, 1761)
Phalaena plebeja Linnaeus, 1761
Phalaena fulvulata Pallas, 1773
Bombyx vidua Fabricius, 1775 (nec Poda, 1763; junior primary homonym)
Phalaena glaucescens Goeze, 1781
Phalaena glaucofasciata Goeze, 1781
ab. brunnea Closs, xxxx
ab. extrema Rebel, xxxx
ab. flava Wood, xxxx
ab. nigra Tutt, xxxx
ab. nigrobasalis Spuler, xxxx
ab. obscura Prout, xxxx
ab. passetii Thierry-Mieg, xxxx
ab. luteata Hennin, 1910
plebeia Prout, 1912 (unjustified emendation of plebeja)
ab. dealbata Klemensiewicz, 1913
ab. dilutior Heinrich, 1916
ab. unicolor Heinrich, 1916
ab. szymanskii Isaak, 1920
ab. muliercula Stephan, 1923
ab. extremata Buresch, 1924 (misspelling of extrema?)
ab. contrasta Lempke, 1949
ab. fasciata Lempke, 1949
ab. intermedia Lempke, 1949
ab. variegata Lempke, 1949
ab. albofasciata Cockayne, 1952
ab. cuprea Cockayne, 1952
ab. indigena auct? (Lambillion? Haverkampf?)
Archiearis parthenias fulvata auct. (misspelling of fulvulata)
passeti auct. (Lhomme, 1935) misspelling
Identification
Forewing length: 16-19 mm, males usually slightly smaller than the females. Similar to Archiearis notha (see there for details).
Special characters in the larva: in contrast to the larvae of higher geometrid moths, the larvae of the present species retain all four proleg pairs. However, the prolegs on abdominal segments 3-5 (especially those on segments 3 and 4) are shorter than the proleg pair on abdominal segment 6, thus illustrating an evolutionary intermediate between the fully developed prolegs in the lepidopteran ground plan and the fully reduced prolegs on abdominal segments 3-5 in the higher Geometridae.
Distribution
Widely distributed in Europe and Asia. In Germany recorded from all Bundesländer, but usually not a common species.
Biology
The larvae feed on Birch, mainly Betula pendula and probably also Betula pubescens, and there also are reports of larvae feeding on Rowan (Sorbus aucuparia). Reports of larvae feeding on Beech (Fagus sylvatica) are unconfirmed.
The imagines feed on nectar (predominantly of Willow catkins) and also are attracted to puddles, mud and animal feces (of horses, cows and dogs).
The species prefers deciduous woodland that offers plenty of both birch and willow. Apart from riparian forests, and peatlands, the species is can also be encountered in parks and gardens.
The imagines fly from February until April and are active during the day. They often fly in the direct sunlight in a swarm around tree tops at midday. This might be a kind of courtship behavior, because they fly lower as the afternoon progresses and finally pairs in copula settle on or near the ground towards the evening. The females deposit the eggs in small groups on birch twigs. They show a very characteristic behavior to determine the ideal location for egg deposition. They creep backwards along a twig until their abdomen touches a bud or a branch. At this location they deposit ca. 5-10 eggs in the angle between the twig and the bud/branch. The larvae hatch after one or a few weeks after egg deposition (depending on the temperature) between March and May. During the night they feed on catkins and young leaves, protected by a light silk web. Often the larvae also use silk to spin together two leaves as a hideout during the day when they do not feed. For pupation the larvae hide beneath soft bark or bore into rotten wood, pupate and overwinter as a pupa. The imagines may eclose in the following spring or a year later.
Contact

Zoographia Germaniae is authored and maintained by Niko Prpic-Schäper.
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